Variant report

Variant	nsv613519
Chromosome Location	chr9:11969789-12384802
allele	n/a
Outlinks	Ensembl UCSC

Chromatin state
Related regulatory elements
Target genes
Other information

TF binding
region (count:699)
CpG islands
(count:0)
Chromatin interactive
region (count:10)
LncRNA
region (count:12)
Mature miRNA
region (count: 0)
miRNA target
sites (count:0)

(count:699 , 50 per page) page: 1 2 3 4 5 6 7 ... 14

No.	Transcrition factor	Chromosome Location	Cell Line	Cell type	Cell Stage	Matched TF binding sites
1	ARID3A	chr9:12328968-12329168	HepG2	liver:	n/a	n/a
2	BACH1	chr9:12292536-12292567	H1-hESC	embryonic stem cell:	n/a	n/a
3	BACH1	chr9:12170667-12170713	H1-hESC	embryonic stem cell:	n/a	n/a
4	BATF	chr9:12230772-12231081	GM12878	blood:	n/a	n/a
5	BATF	chr9:12329455-12329670	GM12878	blood:	n/a	n/a
6	BCLAF1	chr9:12329426-12329668	GM12878	blood:	n/a	n/a
7	BHLHE40	chr9:12328995-12329158	K562	blood:	n/a	n/a
8	BHLHE40	chr9:12329021-12329257	GM12878	blood:	n/a	n/a
9	BHLHE40	chr9:12328884-12329725	HepG2	liver:	n/a	n/a
10	BRCA1	chr9:12368889-12368916	GM12878	blood:	n/a	n/a
11	BRCA1	chr9:12011400-12011402	GM12878	blood:	n/a	n/a
12	CBX3	chr9:12054451-12054644	K562	blood:	n/a	n/a
13	CEBPB	chr9:12129071-12129335	HepG2	liver:	n/a	n/a
14	CEBPB	chr9:12193945-12194181	A549	lung:	n/a	chr9:12194071-12194082 chr9:12194071-12194084
15	CEBPB	chr9:12119993-12120079	A549	lung:	n/a	chr9:12120059-12120068 chr9:12120057-12120070 chr9:12120057-12120068
16	CEBPB	chr9:12180768-12180970	IMR90	lung:	n/a	n/a
17	CEBPB	chr9:12208833-12209071	HepG2	liver:	n/a	chr9:12208898-12208909 chr9:12208899-12208908 chr9:12208897-12208910
18	CEBPB	chr9:12367335-12367535	HepG2	liver:	n/a	chr9:12367443-12367454
19	CEBPB	chr9:12195456-12195797	HepG2	liver:	n/a	chr9:12195611-12195622
20	CEBPB	chr9:12195488-12195695	K562	blood:	n/a	chr9:12195611-12195622
21	CEBPB	chr9:12288039-12288167	HepG2	liver:	n/a	n/a
22	CEBPB	chr9:12012267-12012412	A549	lung:	n/a	n/a
23	CEBPB	chr9:12195471-12195788	IMR90	lung:	n/a	chr9:12195611-12195622
24	CEBPB	chr9:12195470-12195708	H1-hESC	embryonic stem cell:	n/a	chr9:12195611-12195622
25	CHD1	chr9:12314638-12314797	GM12878	blood:	n/a	n/a
26	CHD2	chr9:12314733-12314838	GM12878	blood:	n/a	n/a
27	CHD2	chr9:12319226-12319370	GM12878	blood:	n/a	n/a
28	CTCF	chr9:12142395-12142451	LNCaP	prostate:	n/a	n/a
29	CTCF	chr9:12329500-12329650	AG10803	skin:	n/a	n/a
30	CTCF	chr9:12329480-12329630	NHEK	skin:	n/a	n/a
31	CTCF	chr9:12329422-12329710	LNCaP	prostate:	n/a	n/a
32	CTCF	chr9:12329480-12329630	AG09319	gingival:	n/a	n/a
33	CTCF	chr9:12231143-12231170	GM20000	blood:	n/a	n/a
34	CTCF	chr9:12329500-12329650	GM12874	blood:	n/a	n/a
35	CTCF	chr9:12329440-12329590	WI-38	lung:	n/a	n/a
36	CTCF	chr9:12329400-12329550	HFF-Myc	foreskin:	n/a	n/a
37	CTCF	chr9:12328760-12328910	RPTEC	kidney:	n/a	n/a
38	CTCF	chr9:12329423-12329765	A549	lung:	n/a	n/a
39	CTCF	chr9:12329413-12329713	GM12892	blood:	n/a	n/a
40	CTCF	chr9:12219074-12219108	GM10248	blood:	n/a	n/a
41	CTCF	chr9:12329500-12329650	BJ	skin:	n/a	n/a
42	CTCF	chr9:12329500-12329650	AG04449	skin:	n/a	n/a
43	CTCF	chr9:12055107-12055140	GM13976	blood:	n/a	n/a
44	CTCF	chr9:12329500-12329650	GM12868	blood:	n/a	n/a
45	CTCF	chr9:12329500-12329650	HA-sp	spinal cord:	n/a	n/a
46	CTCF	chr9:12329442-12329697	Medullo	brain:	n/a	n/a
47	CTCF	chr9:12329309-12329662	MCF-7	breast:	n/a	n/a
48	CTCF	chr9:12329820-12329970	AG04450	lung:	n/a	n/a
49	CTCF	chr9:12329520-12329670	Hela-S3	cervix:	n/a	n/a
50	CTCF	chr9:12329462-12329694	GM13976	blood:	n/a	n/a

No data

(count:10 , 50 per page) page: 1

No.	Distal block	Cell Line	Cell type
1	chr18:21570976..21573123-chr9:12145530..12147030,2	MCF-7	breast:
2	chr9:12182493..12185046-chr9:12186407..12189303,2	K562	blood:
3	chr9:12094328..12096426-chr9:12096756..12098736,2	K562	blood:
4	chr9:12182493..12185046-chr9:12186407..12189303,2	K562	blood:
5	chr16:89002000..89004763-chr9:12244429..12246932,2	MCF-7	breast:
6	chr9:12094328..12096426-chr9:12096756..12098736,2	K562	blood:
7	chr9:12268631..12271178-chr9:12273434..12276211,2	K562	blood:
8	chr9:12268631..12271178-chr9:12273434..12276211,2	K562	blood:
9	chr9:10365979..10366914-chr9:12329143..12329893,2	MCF-7	breast:
10	chr9:10855828..10856443-chr9:12329128..12329639,2	MCF-7	breast:

(count:12 , 50 per page) page: 1

No.	lncRNA name	Chromosome Location	lncRNA alias
1	lnc-MPDZ-9	chr9:12106821-12106865	NONHSAT130219
2	lnc-TYRP1-4	chr9:12287320-12288238	NONHSAT130222
3	lnc-MPDZ-9	chr9:12136273-12136401	NONHSAT130219
4	lnc-MPDZ-9	chr9:12088660-12089015	NONHSAT130219
5	lnc-MPDZ-2	chr9:12139007-12139044	XLOC_007648
6	lnc-MPDZ-2	chr9:12159117-12159147	XLOC_007648
7	lnc-MPDZ-2	chr9:12098714-12099015	XLOC_007648
8	lnc-MPDZ-2	chr9:12116821-12116865	XLOC_007648
9	lnc-MPDZ-2	chr9:12098660-12099015	XLOC_007648
10	lnc-MPDZ-2	chr9:12116821-12116865	XLOC_007648
11	lnc-MPDZ-9	chr9:12149117-12149147	NONHSAT130219
12	lnc-MPDZ-2	chr9:12146273-12146401	XLOC_007648

No data

Variant related genes	Relation type
ENSG00000231491	TF binding region
ENSG00000224935	TF binding region
RNU2-47P	TF binding region
ENSG00000261226	chromatin interactions
ENSG00000168234	chromatin interactions

Extended variants
information (count: 6034 )
Associated
traits (count: 111 )

Variant overlapped rSNPs/rCNVs (count:6034 , 50 per page) page: 1 2 3 4 5 6 7 ... 121

No.	Variant name	Chromosome position	Chromatin state	Related regulatory elements	Target genes	Extended variants	Associated traits
1	rs549716331	chr9:11972811-11972812	Enhancers	n/a	n/a	Overlapped CNVs	n/a
2	rs189027366	chr9:11972821-11972822	Enhancers	n/a	n/a	Overlapped CNVs	n/a
3	rs78897159	chr9:11972864-11972865	Enhancers	n/a	n/a	Overlapped CNVs	n/a
4	rs554781765	chr9:11972926-11972927	Enhancers	n/a	n/a	Overlapped CNVs	n/a
5	rs568137692	chr9:11972927-11972928	Enhancers	n/a	n/a	Overlapped CNVs	n/a
6	rs533957171	chr9:11972950-11972951	Enhancers	n/a	n/a	Overlapped CNVs	n/a
7	rs566629462	chr9:11972953-11972954	Enhancers	n/a	n/a	Overlapped CNVs	n/a
8	rs149029538	chr9:11972954-11972955	Enhancers	n/a	n/a	Overlapped CNVs	n/a
9	rs62551976	chr9:11972959-11972960	Enhancers	n/a	n/a	LD-proxies of rSNP Overlapped rCNV	n/a
10	rs372066640	chr9:11972982-11972983	Enhancers	n/a	n/a	Overlapped CNVs	n/a
11	rs556187966	chr9:11973019-11973020	Enhancers	n/a	n/a	Overlapped CNVs	n/a
12	rs539486706	chr9:11973059-11973060	Enhancers	n/a	n/a	Overlapped CNVs	n/a
13	rs576709278	chr9:11973070-11973071	Enhancers	n/a	n/a	Overlapped CNVs	n/a
14	rs143010739	chr9:11973093-11973094	Enhancers	n/a	n/a	Overlapped CNVs	n/a
15	rs139422341	chr9:11973100-11973101	Enhancers	n/a	n/a	Overlapped CNVs	n/a
16	rs148279320	chr9:11973110-11973111	Enhancers	n/a	n/a	Overlapped CNVs	n/a
17	rs115098747	chr9:11973122-11973123	Enhancers	n/a	n/a	Overlapped CNVs	n/a
18	rs141298851	chr9:11973186-11973187	Enhancers	n/a	n/a	Overlapped CNVs	n/a
19	rs541185993	chr9:11973199-11973200	Enhancers	n/a	n/a	Overlapped CNVs	n/a
20	rs570099638	chr9:11975002-11975003	Enhancers	n/a	n/a	Overlapped CNVs	n/a
21	rs533261519	chr9:11975033-11975034	Enhancers	n/a	n/a	Overlapped CNVs	n/a
22	rs543292357	chr9:11975036-11975037	Enhancers	n/a	n/a	Overlapped CNVs	n/a
23	rs146860989	chr9:11975042-11975043	Enhancers	n/a	n/a	Overlapped CNVs	n/a
24	rs370787481	chr9:11975049-11975050	Enhancers	n/a	n/a	Overlapped CNVs	n/a
25	rs529005143	chr9:11975112-11975113	Enhancers	n/a	n/a	Overlapped CNVs	n/a
26	rs529737729	chr9:11975139-11975140	Enhancers	n/a	n/a	Overlapped CNVs	n/a
27	rs549082200	chr9:11975159-11975160	Enhancers	n/a	n/a	Overlapped CNVs	n/a
28	rs549154205	chr9:11975187-11975188	Enhancers	n/a	n/a	Overlapped CNVs	n/a
29	rs565399129	chr9:11975192-11975193	Enhancers	n/a	n/a	Overlapped CNVs	n/a
30	rs73402806	chr9:11975193-11975194	Enhancers	n/a	n/a	LD-proxies of rSNP Overlapped rCNV	n/a
31	rs550961669	chr9:11975195-11975196	Enhancers	n/a	n/a	Overlapped CNVs	n/a
32	rs77559688	chr9:11975605-11975606	Enhancers	n/a	n/a	Overlapped CNVs	n/a
33	rs192329221	chr9:11975650-11975651	Enhancers	n/a	n/a	Overlapped CNVs	n/a
34	rs140764166	chr9:11975709-11975710	Enhancers	n/a	n/a	Overlapped CNVs	n/a
35	rs150089318	chr9:11975738-11975739	Enhancers	n/a	n/a	Overlapped CNVs	n/a
36	rs528167819	chr9:11975743-11975744	Enhancers	n/a	n/a	Overlapped CNVs	n/a
37	rs375322554	chr9:11975759-11975760	Enhancers	n/a	n/a	Overlapped CNVs	n/a
38	rs138359747	chr9:11975777-11975778	Enhancers	n/a	n/a	Overlapped CNVs	n/a
39	rs570890345	chr9:11975788-11975789	Enhancers	n/a	n/a	Overlapped CNVs	n/a
40	rs73402811	chr9:11975795-11975796	Enhancers	n/a	n/a	LD-proxies of rSNP Overlapped rCNV	n/a
41	rs182946807	chr9:11975799-11975800	Enhancers	n/a	n/a	Overlapped CNVs	n/a
42	rs188022507	chr9:11975815-11975816	Enhancers	n/a	n/a	Overlapped CNVs	n/a
43	rs147009642	chr9:11975821-11975822	Enhancers	n/a	n/a	Overlapped CNVs	n/a
44	rs367629402	chr9:11975835-11975836	Enhancers	n/a	n/a	Overlapped CNVs	n/a
45	rs575814519	chr9:11975841-11975842	Enhancers	n/a	n/a	Overlapped CNVs	n/a
46	rs114791891	chr9:11975847-11975848	Enhancers	n/a	n/a	Overlapped CNVs	n/a
47	rs567009814	chr9:11975856-11975857	Enhancers	n/a	n/a	Overlapped CNVs	n/a
48	rs193266910	chr9:11975861-11975862	Enhancers	n/a	n/a	Overlapped CNVs	n/a
49	rs371892008	chr9:11975897-11975898	Enhancers	n/a	n/a	Overlapped CNVs	n/a
50	rs575339554	chr9:11975910-11975911	Enhancers	n/a	n/a	Overlapped CNVs	n/a

Variant related diseases (count:111)

Disease	PMID	Source
Disorders of sex development	21048976	CNVD
Honadal dysgenesis	21048976	CNVD
Medulloblastoma	21979893	CNVD
Lung cancer	16740712	CNVD
Malignant peripheral nerve sheath tumor	18522746	CNVD
Basal cell lymphoma	17170743	CNVD
Follicular lymphoma	17170743	CNVD
Oral squamous cell carcinoma	17134496	CNVD
Chronic lymphocytic leukemia	21546498	CNVD
abnormal development	18461090	CNVD
Glioblastoma multiforme	21080181	CNVD
Breast cancer	16608533	CNVD
Hodgkin''s lymphoma	18641027	CNVD
Acute lymphoblastic leukemia	20067559	CNVD
Breast cancer	16272173	CNVD
Hepatocellular carcinoma	16750200	CNVD
Breast cancer	17133270	CNVD
Endometrioid adenocarcinoma	16974079	CNVD
Cervical cancer	21063398	CNVD
Retinoblastoma	22278416	CNVD
Splenic marginal zone lymphoma	18492102	CNVD
Melanoma	18172304	CNVD
Biliary cancer	19435499	CNVD
Gastric cancer	21586687	CNVD
Esophageal squamous carcinoma	18350619	CNVD
Myopathy	18421352	CNVD
Neuroblastoma	18664255	CNVD
Acute lymphoblastic leukemia	18957548	CNVD
Myeloproliferative neoplasm	17564968	CNVD
Malignant meningioma	17937814	CNVD
Glioma	21971842	CNVD
Esophageal squamous carcinoma	21637470	CNVD
Ewing''s sarcoma	21437220	CNVD
Non-syndromic sensorineural hearing loss	18471307	CNVD
Astrocytoma	17387387	CNVD
Breast cancer	17603634	CNVD
Endometrial cancer	22040021	CNVD
Breast cancer	21264507	CNVD
Glioblastoma multiforme	19960244	CNVD
Glioblastoma multiforme	17387387	CNVD
Hepatocellular carcinoma	22174799	CNVD
Oligodendroglial tumors	17285580	CNVD
Cutaneous squamous cell carcinomas	19131950	CNVD
Small bowel adenocarcinoma	21586687	CNVD
Basal cell lymphoma	16317097	CNVD
Diffuse large b-cell lymphoma	16317097	CNVD
Chordoma	18071362	CNVD
Acute lymphoblastic leukemia	22237106	CNVD
Chordoma	21602918	CNVD
Renal cell carcinoma	18592004	CNVD
Developmental delay	19490664	CNVD
Pleomorphic xanthoastrocytoma	20730472	CNVD
microdeletion syndrome	16199537	CNVD
Non-syndromic sensorineural hearing loss	21987784	CNVD
Cancer	20164919	CNVD
Chronic myeloid leukemia	21384125	CNVD
Ovarian cancer	21720365	CNVD
Non-syndromic sensorineural hearing loss	21030649	CNVD
small cell lung cancer	20016488	CNVD
Acute lymphoblastic leukemia	20435627	CNVD
T-cell prolymphocytic leukemia	19278963	CNVD
Lung cancer	18438408	CNVD
Urothelial carcinoma	21177765	CNVD
Oral cancer	21386901	CNVD
lymphocytic leukemia	21291569	CNVD
Acute lymphoblastic leukemia	21339820	CNVD
Intellectual disability	22102821	CNVD
Breast cancer	21785460	CNVD
Esophageal adenocarcinoma	19417022	CNVD
Hodgkin''s lymphoma	17606441	CNVD
Breast cancer	21858162	CNVD
Renal cell carcinoma	21668985	CNVD
Myelodysplastic syndrome	21251322	CNVD
Autism	22495311	CNVD
Hepatocellular carcinoma	18803288	CNVD
Breast cancer	21364760	CNVD
Myelofibrosis	22110671	CNVD
Glioblastoma	19074898	CNVD
Malignant melanoma	19074898	CNVD
Prostate cancer	16573809	CNVD
Metanephric adenoma	20802469	CNVD
Cancer	20164920	CNVD
Developmental delay	21147756	CNVD
small cell lung cancer	21764851	CNVD
Glioblastoma multiforme	22286061	CNVD
Glioma	17123091	CNVD
Lung cancer	16773561	CNVD
Colorectal cancer	16272173	CNVD
Esophageal squamous carcinoma	18405350	CNVD
Gastric cancer	16891809	CNVD
Mental retardation	17847001	CNVD
Cutaneous squamous cell carcinomas	17420988	CNVD
Acute lymphoblastic leukemia	21390130	CNVD
Pilocytic astrocytoma	18670637	CNVD
Acute lymphoblastic leukemia	17690704	CNVD
Acute myeloid leukemia	21251322	CNVD
Breast cancer	21611746	CNVD
Melanoma	22183965	CNVD
low-grade B-cell lymphoma tumor	18367492	CNVD
Type 2 diabetes	21526130	CNVD
Non-syndromic sensorineural hearing loss	17122850	CNVD
Schizophrenia	20967226	CNVD
early-passage human iPS cells	21368824	CNVD
Williams Syndrome	20824207	CNVD
Schizophrenia	23813976	CNVD
Non-syndromic sensorineural hearing loss	18451855	CNVD
Breast cancer	22522925	CNVD
Non-syndromic sensorineural hearing loss	22037309	CNVD
Autism	20858261	CNVD
Epilepsy	20858261	CNVD
Mental retardation	20858261	CNVD

Chromatin state (count:232 , 50 per page) page: 1 2 3 4 5

No.	Chromosome Location	Chromatin state	Cell line	Tissue
1	chr9:11972800-11973200	Enhancers	Placenta Amnion	Placenta Amnion
2	chr9:11975000-11975200	Enhancers	Pancreatic Islets	Pancreatic Islet
3	chr9:11975600-11976200	Enhancers	Fetal Brain Male	brain
4	chr9:11976000-11976200	Enhancers	Pancreatic Islets	Pancreatic Islet
5	chr9:11977400-11977600	Enhancers	Pancreatic Islets	Pancreatic Islet
6	chr9:11991400-11994000	Enhancers	Cortex derived primary cultured neurospheres	brain
7	chr9:11991800-11992200	Enhancers	H9 Derived Neuronal Progenitor Cultured Cells	ES cell derived
8	chr9:11991800-11992200	Enhancers	Fetal Heart	heart
9	chr9:11991800-11993400	Enhancers	Foreskin Melanocyte Primary Cells skin03	Skin
10	chr9:11992400-11993200	Enhancers	Ganglion Eminence derived primary cultured neurospheres	brain
11	chr9:11993000-11993800	Enhancers	hESC Derived CD56+ Ectoderm Cultured Cells	ES cell derived
12	chr9:11995800-11996600	Enhancers	hESC Derived CD184+ Endoderm Cultured Cells	ES cell derived
13	chr9:11995800-11998200	Weak transcription	H1 Derived Neuronal Progenitor Cultured Cells	ES cell derived
14	chr9:11996600-11996800	Enhancers	iPS DF 6.9 Cell Line	embryonic stem cell
15	chr9:12017400-12019600	Weak transcription	Fetal Heart	heart
16	chr9:12019600-12021400	Enhancers	Fetal Heart	heart
17	chr9:12046400-12046800	ZNF genes & repeats	Foreskin Melanocyte Primary Cells skin03	Skin
18	chr9:12046800-12048000	Weak transcription	Foreskin Melanocyte Primary Cells skin03	Skin
19	chr9:12048000-12048400	Enhancers	Foreskin Melanocyte Primary Cells skin03	Skin
20	chr9:12054200-12054600	ZNF genes & repeats	H1 BMP4 Derived Trophoblast Cultured Cells	ES cell derived
21	chr9:12057600-12059000	Enhancers	Foreskin Melanocyte Primary Cells skin03	Skin
22	chr9:12058400-12060400	Enhancers	Fetal Intestine Large	intestine
23	chr9:12059000-12060000	Weak transcription	Foreskin Melanocyte Primary Cells skin03	Skin
24	chr9:12060000-12065800	Enhancers	Foreskin Melanocyte Primary Cells skin03	Skin
25	chr9:12060200-12060400	Enhancers	Foreskin Melanocyte Primary Cells skin01	Skin
26	chr9:12060400-12061400	Weak transcription	Foreskin Melanocyte Primary Cells skin01	Skin
27	chr9:12060400-12065000	Weak transcription	Fetal Intestine Large	intestine
28	chr9:12061400-12061800	Enhancers	Foreskin Melanocyte Primary Cells skin01	Skin
29	chr9:12061400-12062000	Enhancers	Placenta Amnion	Placenta Amnion
30	chr9:12061600-12061800	Enhancers	hESC Derived CD56+ Mesoderm Cultured Cells	ES cell derived
31	chr9:12061800-12064200	Weak transcription	hESC Derived CD56+ Mesoderm Cultured Cells	ES cell derived
32	chr9:12064200-12065600	Enhancers	hESC Derived CD56+ Mesoderm Cultured Cells	ES cell derived
33	chr9:12065000-12065800	Enhancers	Fetal Intestine Large	intestine
34	chr9:12076400-12076600	Flanking Bivalent TSS/Enh	Breast Myoepithelial Primary Cells	Breast
35	chr9:12076400-12076800	Flanking Bivalent TSS/Enh	ES-UCSF4 Cell Line	embryonic stem cell
36	chr9:12076600-12076800	Bivalent/Poised TSS	Breast Myoepithelial Primary Cells	Breast
37	chr9:12091200-12096400	Enhancers	Placenta Amnion	Placenta Amnion
38	chr9:12091600-12092600	Enhancers	H1 BMP4 Derived Trophoblast Cultured Cells	ES cell derived
39	chr9:12091800-12092200	Enhancers	HUES6 Cell Line	embryonic stem cell
40	chr9:12092600-12096200	Weak transcription	H1 BMP4 Derived Trophoblast Cultured Cells	ES cell derived
41	chr9:12096200-12096400	Enhancers	H1 BMP4 Derived Trophoblast Cultured Cells	ES cell derived
42	chr9:12096400-12101600	Weak transcription	H1 BMP4 Derived Trophoblast Cultured Cells	ES cell derived
43	chr9:12101600-12102200	ZNF genes & repeats	ES-UCSF4 Cell Line	embryonic stem cell
44	chr9:12101600-12102400	ZNF genes & repeats	H1 BMP4 Derived Trophoblast Cultured Cells	ES cell derived
45	chr9:12101800-12102800	ZNF genes & repeats	hESC Derived CD184+ Endoderm Cultured Cells	ES cell derived
46	chr9:12102400-12104200	Weak transcription	H1 BMP4 Derived Trophoblast Cultured Cells	ES cell derived
47	chr9:12104200-12104400	ZNF genes & repeats	H1 BMP4 Derived Trophoblast Cultured Cells	ES cell derived
48	chr9:12135800-12137400	Enhancers	Fetal Heart	heart
49	chr9:12137600-12138400	Enhancers	Placenta Amnion	Placenta Amnion
50	chr9:12137800-12138200	Enhancers	H1 BMP4 Derived Trophoblast Cultured Cells	ES cell derived

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